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    15 September 1986, Volume 5 Issue 03
    A lower molar of Pliopithecus from Tongxin, Ningxia Hui Autonomous Region
    Qiu Zhanxiang, Guan Jian
    1986, 5(03):  201-207. 
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    The subject of the present paper, a left M2 of Pliopithecus, BPV 261 (housed in Beijing Natural History Museum) , was purchased, purely by chance, in the early spring of 1985 from a "dragon-bone" digger in situ, where he had successfully found teeth and bones of Platybclodon, Listriodon and Stephanocemas. The locality, called Maerzuizigou, lies near the village Dingjiaergou, which is 18 km northeast to Tongxin, the county center. Its geographie position is indicated in text-figure 1.
    From the same locality, and chiefly frorn the surrounding area, more materials were collected by the field parties of both the Institute of Vertebrate Paleontology and Paleoanthropology and Beijing Natural History Museum during the last several years. Although the study of the materials is still under way and the geological setting of the area concerned is still being prepared, the geological age of the area in general is clear: It is a little older than the typical Tung-gur faunal age, probably comparable to MN 6 or 7 in the European mammalian biochronology.
    During the first half of the present century there were some reports on the possible oceurrence of pliopithecine material in North China. Schlosser (1924) first reported an M3 under the name of Pliopithecus posthumus. However, the tooth is too heavily worn to warrant its taxonomic position as proposed by M. Sehlosser. Recently Ginsburg and Mein (1980) proposed to refer it to a gibbon genus: Krishnapithecus. Later Bohlin (1946) deseribed some very badly preserved specimens and erected for them a new genus, Kansupithecus (without designation of species) , allegedly belonging to the pliopithecine group. Because of their poor state of preservation (a practically edentulous lower jaw fragment and some tooth splits) , again, their true nature is uncertain. Szalay and Delson (1979) provisionally referred them to tPliopithecidae incerta sedis. The Tongxin material, though consisting of only one tooth, represents therefore the first authentie record of the genus in China. In addition, the preservation of the tooth is excellent, the erown features are all clearly demonstrated.
    A cursory comparison is enough to reveal that our Tongxin specimen conforms very well with Hürzeler’s diagrammatie presentation of the pliopithecine lower molars based on G?riach material (Hürzeler, 1954, fig. 14) . The features our M2 has in common with Hürzeler’s diagram are the following: 1. The labial wall of the tooth is marked'y slan- ting, so that the labial eusps approximate the lingual ones strongly at the top of the erown. 2. The trigonid is large. basin-shaped. and blocked posteriorly by a well develop- ed metalophid. 3. The talonid eonsists of three cusps: hypoconid, entoconid and hypoco- nulid, without indication of the sixth cusp. 4. The distance between the hypo- and entoconid is longer than that between the proto- and metaconid. 5. The metastylid is tiny, but still discernible. 6. There is a "hinter Hauptleiste" (Remane's term) connecting the hypoeonulid and entoconid. 7. There is a "pliopithecine triangle", which is typical for all the members of the group. 8. There is a weak connection between the hypoconid and metaconid on the bottom of the talonid. 9. The labial eingulum is well developed, interrupted only at the midpoint. The enumeration of these characters in common leaves no room for doubt that the Tongxin specimen should refer to the genus Pliopithecus.
    The systematies of the pliopithecine group in now rather complicated. Ginshurg and Mein (1980) proposed to split Pliopithecidae into two subfamilies: Crouzelinae and Pliopithecinae. However, Szalay and Delson (1979) already seriously doubted the validily of the genus Crouzelia, which served the base for Ginsburg and Mein's separation of the subfamily Crouzelinae, based chiefly on the assumption that Crouzelia was based on wrong identification of the deciduous teeth as permanent ones. The careful observation of the cast of the type speeimen shows that the original teeth identification was correct, therefore Ginsburg and Mein's subdivision of the family Pliopithecidae is to be considered tenable. Our speeimen from Tongxin should obviously be exeluded from the subfamily Crouzelinae, which is characterized by the lophodont tendeney of the main cusps, reduction of the entoconid and hypoconulid and less developed cingulum. According to Ginsburg and Mein, Pliopithecinae consists of only one genus and three species: Pliopithecus piveteaui, P. antiquus and P. vindobonensis. The Tongxin specimen is evidently closest to the later speeies in both morphology and size. Nevertheless, they differ still considerably: 1. The Tongxin specimen is even larger in size than that in the iater species. In fact our specimen represents the largest individual ever found for the genus Pliopithecus (see table 1) . 2. The secondary structures on the erown surface of the Tongxin specimen are rieher developed. 3. The "pliopithecine triangle"is still not completely closed in our specimen.
    Our new specimen may well represent a new species of the genus, but the paueity of the material made us hesitating in doing so.
    Preliminary research on the fossil gibbon of Pleistocene China
    Cu Yumin
    1986, 5(03):  208-219. 
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    In this paper material from 15 loealities is diseussed. 37 teeth were collected from the Guangsi Zhuang Autonomous Region of China during explorations under the leadership of professor Pei Wencheng in 1956-1957, and 5 of them were provided by Han Defen from Ling-yan cave. Liujiang County, Guangxi. 20 teeth, collected from a medicinal supply company,came from unnamed cave sites.
    There is only one upper premolar.It has two cusps.There is a well-developed cingulum which runs from lingual aspect to the distal and buceal aspects of the tooth.
    There are 30 upper molar specimens, of which 25 are either M1 or M2,5 are M3.For M1 or M2 each has four cusps; the paracone and metacone are highest. followed by the protoeone and hypoeone. In M3 the hypocone is less-developed. In some of the upper molars a well-developed lingual cingulum is expressed on the lingual aspect of the protocone. The size of some specimens which possess cingulum is large, a notch and a ridge are also present on their mesial border. The other teeth possessing cingulum are small and the noteh and ridge are lacking. It is possible that the former are of M2 and the latter are of M1.The cingulum on three specimens of M1 or Ma is not seen and the size of these teeth are smaller than those having cingulum.
    The lower molars possess five cusps:a lypoconulid is added to each molar and it is positioned eentrally. The protoeonid and metaeonid are connected by a eotoeristid. The trigonid basin is narrower than the talonid.
    Frisch (1965) documented that Hylobates concolor is distinguished by well-developed cingulum on the lingual aspect of the upper molars. I therefore deduced that the teeth possessing cingulum, which came from Guangxi, can be attributed to Hylobates concolor. In addition to the black gibbon two extant specics of gibbon(H.lar,H. hoolock) still exist in China. Those teeth without cingulum are possibly attributable to either H. lar or H. hoolock. The teeth of H.lar are smaller than those without cingulum found in Guangxi. As the distribution of this species is limited in our country,it is thought that these teeth with no cingulum possibly cannot belong to H.lar. No trace of cingulum could be found in nearly the total H. hoolock specimens, consequently,I regard those teeth without eingulum found in Guangxi as H. hoolock at present.
    The chronologic distribution of the gibbon in China can be divided into three periods: Recent, Historie (400 B. Y.) and Pleistocene. Historical accounts (Gao Yaoting et al., 1981) and fossil evidenee show that the gibbon once ranged widely in China up to 31°N latitude. More recently, it reachs only to 26°N (see fig.5).
    Cingulum is considered to be a primitive character. Moreover, a difference in the number of chromosomes exists between the three species of extant gibbon in China.The chromosome number in 94% of black gibbons (H. concolor) is 2n=52; in the others(H. lar, H. hoolock) 2n=44 in more than 90% of individuals,H. concolor may therefore be a more conservative animal than the other two species.
    The evolution of the gibbon is still an open question. Recently, new material has been continuously found in China. Since Neogene times the vegetation has been favourable for the gibbon to live. It is possible that the ancestor of the gibbon may be found in China.
    Study of temporal bone of Xujiayao Man
    Wu Maolin
    1986, 5(03):  220-226. 
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    This paper deals with a fossil human temporal from Xujiayao site in 1979 where previously 17 human fossils were found.
    It has many characters identical to those of early Homo sapiens or intermediate between Homo erectus and modern man. The features include: the robustness of the supramastoid erest, the postglenoid process, the position and the orientation of the tympanic plate, the position of the tympanomastoid fissure, the orientation of the pyramid axis, the distance between the styloid process and the posterior wall of the porus, the entire mastoid process deviates slightly medianwards, ete.
    The temporal bone of Xujiayao possesses some features as seen in modern man such as the form of the squama, the position of the supramastoid erest, the form of tympanic plate, the styloid process, tho foramen stylomastoideum and the supramastoid erest arranged in a straight line, the sigmoid suleus, ete.
    There are some features close to those of Peking Man such as absence of true articular tuberele, the thickness of the mastoid portion, ete.
    Anthropological characters of the human crania from Loulan Site Xinjiang
    Han Kangxin
    1986, 5(03):  227-242. 
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    Six skulls from an ancient cemetery at Loulan site in 1980 are studied in this paper. The age of this cemetery, according to C4 dating, is about 2000 B. P. Among six observed skulls four belong to males and two females.
    The morphological observation and measurements show that five skulls have Europoid's characters. These skulls mainly represent the natives of Europoid in the region, and their physical characters are mostly similar to that of Indo-Afghan pattern. One Mongoloid eranium is found in the cemetery. It is suggested that the ethnic elements of Loulan Kingdom did not consist of single race.
    In the comparison with other ancient eranial series of Central Asia, Loulan's erania are more similar to that of Saka in Pamir morphologically, whereas, they are not homogeneous with the brachyeephalic Saka-Wusun's erania in Tianshan-Alei and Zhaosu and are close to that of Pamir's Saka in anthropological relationship. These Loulan's natives had exchanged with the population of Han Dynasty in culture from the archaeoIogic evidences.
    Secular change and geographical variation in Chinese Neolithic and modren inhabitants: A statistical study of cranial metric traits
    Wang Linghong
    1986, 5(03):  243-258. 
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    China is rich in Neolithie human skeletons as well as Paleolithic human remains. The former is even more abundant in number. Nevertheless, the evolutionary aspeets of these skeletons and their morphological differences from modern ones have not drawn sufficient attention. The present paper aims to examine the nature of morphologieal evolution in Holocene human populations of China. In literature were described about 20 Chinese Neolithie sites yielding more or less human crania, at most 30—40, at least 1—2. Among them, southern and northen populations were separated, and the North China group was further subdivided into eastern and western subgroups.
    In order to involve as many groups and variables as possible, two techniques were used for preparing the data-sets. These are 1)combination of sample groups and 2) estimation of missing values.
    There are two reasons for eombining sample groups. First of all, in some cases, there were more than one group sampled from the same population. Usually, one can not know which is more representative. Combining them is a better way than choosing at random, as long as there exists reasonable similarity among the samples. Secondly, combining is useful when any sample group does not contain sufficient number of samples, or has all variables represented. The member groups can be mutually complementary.
    In this study, after combining samples there were still some missing values in the data-sets. And these missing values involved such important groups as the Zengpiyan group and such important variables as eranial base length and facial length. In order to maintain both these groups and these variables in the statistical study, the missing values had to be estimated. Stepwise regression analysis was used for that. The estimated values have been proved reliable by both the F-test and the correlation test.
    Cluster analysis, factor analysis, diseriminant analysis and two way variance analysis were used in this study. The results produced by these four techniques are quite consistent.
    1. Cluster analysisIn the dendrogram of the male groups based on Penrose's shape distances, Neolithic north and south groups, and modern north and south groups respectively form their own blocks. One exception is the Gansu-Henan Neolithie group, which was excavated by the Swedish geologist and archaeologist J. Andersson and studied by the Canadian anthropologist D. Blaek. The two regional branehes of the same period join with each other respeetively. Then, the Neolithie branch and the modern branch link at a higher level (greater distance) . This shows that there is a bigger difference in terms of time than of region. In other words, the secular change is generally more significant than the geographical variation. In addition, in the Neolithic branch and the modern branch, the artificially deformed groups are specially separated as expeeted.
    2. Factor analyssIn the factor analyses of both the male and the female groups, almost all of the linear variables have large positive loadings for thefirst factor, especially facial length, bizygomatie breadth, eranial base length, basi-bregmatie height, minimum frontal breadth and nasal breadth. The first factor, therefore, can be called the size factor.
    For the second factor, upper facial height and nasal height have large loadings. This is the facial height factor.
    The coordinates of the first two factors marking the positions of the groups studied produced the same results as shown above. The two axes distinguish temporal and regional groups respectively. The first factor (X-axis) distinguishes between Neolithic and modern populations. The second factor (Y-axis) distinguishes between north and south populations. Once again, the Gansu-Henan Neolithie group is the only exception.
    The aceuracy of the ehronology for at least some of the samples of the Gansu-Henan group has been questioned based on a review of the Neolithie cultural context. The above analyses support this doubt.
    To compare the male and the female Hemudu specimens is also interesting. Each is represented by only one sample. The male sample (M23 is bigger than the other male groups on the average, while the female one (M 17) is smaller than the other female groups. It is not likely that the tremendous differences of these two samples reflect a extremely big sexual dimorphism of the population. They seem to be merely extreme samples in their respective sexes.
    It is not proper to include the Gansu-Henan group and the Hemudu specimens in the further analyses.
    3. Discriminant analysisThe purpose of the discriminant analyses is to detect the significance of the difference between the temporal groups (i. e. Neolithic and modern group) or the regional groups (i. e. north and south group) in each variables. The analyses of the male groups obtained exactly the same results as did the faotor analysis: there were very significant temporal differences in eranial base length, facial length, basi-bregmatie height, nasal breadth, bizygomatic breadth and minimum frontal breadth and very significant or significant regional differences in nasal height and upper facial height.
    4. Two way variance analysisThe time sequence consists of early Neolithic, late Neolithie and recent times. The region sequence consists of northern, central-eastern and southern China. 9 male groups were chosen to fill in the tablet of the two sequences. The analyses of 8 variables show somewhat gradual changes over time in basi-bregmatic height, nasal breadth, minimum frontal breadth, eranial length and bizygomatic breadth, and somewhat gradual changes over region in upper facial height, nasal height and eranial length.
    Orbital dimensions were not included in the above analyses, because it was noticed that significant errors had been made in measuring orbital height and breadth, and because not all groups contained the mean values of these two measurements. To test whether these two variables are still useful in further discussion, another seattergram was drawn with two axes representing orbital height and orbital breadth respectively. It is shown that there exist differences among the four bloeks: the Neolithic groups have wider and lower orbitals than the modern ones, while the northern groups have higher orbitals than the southern ones. This indieates that errors of measurement of the orbital dimensions do not affeet comparative study to sueh an extent as to obseure the differences in question over time and region.
    Based on the above analyses, the secular change and the geographical variation in question can be summarized as follows.
    Chinese Neolithic residents had larger erania than the modern population. This was reflected in cranial base length, facial length, eranial height, nasal breadth, bizygomatic breadth, minimum frontal breadth and orbital breadth. Their upper facial height and orbital height were slightly lower than in the modern population.
    As to geographical variation, in both Neolithic and recent times northern Chinese inhabitants have higher upper facial height, orbital height and nasal height than southern Chinese ones.
    Generally speaking, the seeular ehange is more significant than the geographical variation.
    The present study is an attempt to consider seeular change and geographical variation at the same time. Obviously, this is often necessary. For instance, sereral Neolithic groups of Shaanxi Yang-shao Culture were characterized by relatively low facial height and relatively prominent prognathism ete. At first sight, they seemed to represent anomalous geographical variation. Taking secular change into account, however, one finds a somewhat inereasing tendency toward facial height and a somewhat decreasing tendeney to prognathism. Therefore, it is not reasonable to regard this Neolithic population as a separate variant of the southern Mongoloid racial stock. The statistical analyses illustrate that to study these two aspects of changes of populations at the same time is not only necessary but also possible if only these changes happened in different eharacters.
    The statistical analyses were carried out under the instruction of Prof. Kazuro Hanihara during the author's stay in the University of Tokyo as visiting scholar. Gratitude is expressed to him and his colleagues, especially Dr. Makiko Kouchi and Mr. Kiyotaka Koizumi for their help. Thanks are also due to Prof. Wu Rukang for his encouragement throughout this study.
    Bone artifacts, ornaments from Xiaogushan Site of Haicheng, Liaoning Province
    Huang Weiwen, Zhang Zhenhong, Fu Renyi, Chen Baofeng, Lin Jingyu, Zhu Mingye, Wu Hongkuan
    1986, 5(03):  259-266. 
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    During the excavation of 1983, a number of bone artifacts and ornaments were discovered in company with mammalian fossils, human fossils and stone artifacts from the Paleolithic layers of Xiaogushan site. The brief deseription is given /below:
    1. Harpoon
    A harpoon with two rows of barbs was probably made of a cannon bone of Artio dactyla. It has a oblate-tapering point and a wedge-shaped base. The shaft is pentagon in cross section. The surface of the shaft shows numerous fine longitudinal striae whieh suggest it had been seraped into shape.
    Such a bone harpoon was not known to have oecurred in China before Neolithie time. It is precisely of the same character as some of the harpoons from the late Magdalenian of Europe.
    2. Javelin point
    A broken javelin point was made of the limb bone of an animal. Its surface is smooth and the eross section is oblatetapering. So far, similar speeimen has never been recorded from Paleolithie China.
    3. Bone needles
    Three bone needles described here are fine ones. Two of them were probably made of ivory and the third one was made of an animal bone. The surfaces are smooth. The shafts are curved and somewhat elliptical in section. The tips well rounded and sharply pointed. The eyes on two specimens are nearly eircular and made by drilling on two faces. The eye on the third one is not completely made.
    In China, such bone needle was only known from Upper Cave of Zhoukoudian. a Late Paleolithic site. The specimen of it is broken just at the eye. Judging from the remains, the eye was not made by drilling but by exeavating or seratehing with a pointed implement.
    4. Perforated teeth
    Four perforated teeth deseribed in this paper were made of canines of small Carnivora or deer, one of them is only a piece of root. All of the holes are loeated at the root of the teeth and made by three different ways: drilling, seratehing and scratehingdrilling. The root was flated by grinding before making the hole.
    Compared with similar specimens of Upper Cave where 125 perforated teeth were found, the perforated teeth from Xiaogushan show a more advanced technique in making the hole. Not a single hole of specimen from Upper Cave might be attributed to drilling.
    5. Perforated shell
    A broken perforated shell was found fron the site. Its outline looks like a coin if preserved well. A hole in the centre was preduced by drilling on two faces. There is a cirele of radial cutting-fissures in its periphery. The red color is still visible from the fissures.
    Among the bone artifacts and ornaments mentioned above, most of them came from Layer 3 of the site, and the remaining came from Layer 2 under Layer 3. The age of the two layers can be attributed to Late Pleistocene based on mammalian fauna. Their absolute dating is from about 40, 000 to 20, 000 y. BP by radiocarbon and thermoluminescence methods.
    A late Paleolithic shell tool from Xinjiang, Shanxi
    Huang Wanbo, Ji Hongxiang, Lin Yongmao
    1986, 5(03):  267-270. 
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    A shell tool associated with mammalian fossils was discovered from the Terrace III of Fen He at Shanjiadian village, xinjiang county (111°15'E; 35°45'N) , Shanxi Pro- vince in 1985. They were unearthed from the sand-gravel deposits under loess. The mammalian remains include Palacoloxodon namadicus, P. cf. takungai, Equus ef. prje- walsh'yi, Sus sp. , Sinomegaceros sp. ete. Considering its stratigraphical position and the fossil assemblage, this site should be dated Late Pleistocene.
    A dermatoglyphic study on population of Han Nationality——Determination of normal values of 7 dactylographic parameters
    Lin Chiping
    1986, 5(03):  271-293. 
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    This paper reports the normal values of 7 dactylographie parameters in population of Han nationality. The sample comprises 4300 males and females. The sbserved and expected frequencies of the 1001 different combinations of ten finger prints of the same individual were given. It proves that the differences of the dactylographie combinations of the different races are more than those of different nationalities of the same race.
    Dermatoglyphics in the patients of leukemia
    Chen Zufen, Zhang Guiru, Bian Shizhong
    1986, 5(03):  294-298. 
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    The dermatoglyphies of i70 leukemie patients have been studied and eompared with 180 eontrols. It was found that Sydney lines of right hands were presented in 8. 81 3. 44% aeute lymphatie leukemia patients and only in 1. 87+0. 92% controls. The patients with C'M[. showed that the rate of simian line of the palmar was inereased. The indes of areh and whorl of the fingers is more higher in the patients of aeute leakemia than the normal. The rate of the palmar white line in leukemie patients is higher (64. 18±4. 14%) than the controls (38+3. 96%. P<0. 01) . The widest point of the major three palmar ereases is positive (>2. 5 mm) , whieh is more frequent than the eontrols (P< 0. 01) . The tiny changs of dermatoglyphies of the leukemie patients showed that the genetie faetors may have an important role in the leukemie pathogenesis.
    Palmar ridge counts in familial deafness
    Neelam Gaur, P.K. Datta
    1986, 5(03):  299-300. 
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    The present study was undertaken to measure a-b, a-d and td ridge counts of deaf patients and to compare the dermatoglyphic differences among deaf individuals. The data consists of 208 heredo-familial Punjabi Khatri and Arora deaf subjects, including 118 males and 90 femeles. The control sample consists of 228 Punjabis, comprising of 128 males and 100 females. The present study indicates lower incidence of 'a-b', 'a-d' and 'td' ridge counts in deaf groups when compared to the corresponding normal groups.