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    Exploration of "behavioral modernity” and “modern human behavior” in human evolution
    YANG Shixia
    Acta Anthropologica Sinica    2025, 44 (02): 181-192.   DOI: 10.16359/j.1000-3193/AAS.2025.0014
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    The evolution and widespread of Homo sapiens is a key event in the human evolution history. By the end of the Late Pleistocene, our species was distributed across every continent except Antarctica, and adopted to varied ecological niches. The behavioral complexity and the cultural diversity are believed to act as one of the most pronounced abilities to get success. The fossil and genetic evidence were applied to discuss the extinction of archaic populations, such as the Denisovans and the Neanderthals and their complicated interbreeding with Homo sapiens. The archaeological records served on the discussion about the behavioral evolution from the emergence of “behavioral complexity or modernity’ to the fully developed modern human ‘behavioral package’. In this paper, we reviewed the discussion on the modern human behavioral evolution in the western side of the Old World. Based on the African archaeological records, the behavioral origins of Homo sapiens can be traced back to the first material culture produced by our species in Africa and phased as Middle Stone Age (MSA); while the western Eurasian records the behavioral complexity raised with the origin of Neanderthals and phased as Middle Paleolithic (MP). Then around the middle of Late Pleistocene, as the wide distribution of Homo sapiens and the extinction of archaic populations, a complete ‘modernity package’ finally developed and shows more cultural diversity globally, which including the advanced lithic tools (blade, microblade), bone tools, increased geographic range, specialized hunting, the use of aquatic resources, long distance exchange, systematic processing and use of pigment, and art and decoration.

    However, recently in Eastern Asia efforts were made to explore the complexity of the ‘behavioral complexity or modernity’, the technological innovations and behavioral adaptations process are still barely known. The lithic assemblages have long been regarded as simple, conservative technologies, and major changes in raw material procurement, core reduction, retouch and typology tended to be clustered until 40 kaBP, without a ‘middle phase’ showing distinguished transition. These years, facing with the high degree of hominin morphological variability which has been suggested by recent fossil and genetic evidence, such as the early modern humans, Denisovans, H. longi, and some other unassigned taxons (Xujiayao and Xuchang), and growing new archeological data, we need to rethink the ‘behavioral modernity’ process happened in Eastern Aisa and its application to the evolutionary history of our species worldwide.

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    Taphonomy of the animal remains from the Shiyu site, Shanxi
    WANG Ying, ZHANG Yue, YANG Shixia, ZHANG Shuangquan
    Acta Anthropologica Sinica    2025, 44 (02): 255-269.   DOI: 10.16359/j.1000-3193/AAS.2024.0098
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    The Shiyu site, situated on the western edge of the Nihewan basin (sensu lato) is an important Late Pleistocene archaeological location in northern China. From an extensive excavation in the last century, a substantial number of lithics, animal bones, and the traces of human fire use were unearthed, along with a limited number of modern human fossils. Recent Optically Stimulated Luminescence (OSL) and Accelerator Mass Spectrometry (AMS) 14C dating have recalibrated the cultural layer to approximately 44.6±1.2 kaBP. The presence of artifacts with some characteristics of the Initial Upper Paleolithic (IUP) assemblage positions Shiyu as a crucial site for understanding early modern human activities in Northeast Asia.

    In this study, 152 faunal bone fragments curated at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) were analyzed from a perspective of vertebrate taphonomy, including age profile assessment, skeletal elements frequency quantification, bone breakage pattern analysis and bone surface modification observation. The objective of this study is to evaluate and discuss the subsistence strategies employed by Shiyu hominins in their exploitation of animal resources. Given the limited sample size, the following tentative conclusions were derived from this research.

    Most notably, bone surface modifications revealed a relatively high intensity of human activity, evidenced by a significant frequency of human-inflict damages on bones, including cut marks, percussion marks, and percussion notches. In contrast, natural modifications such as carnivore or rodent tooth marks, sedimentary abrasions, and polish were minimal. Even when considering the potential biases in specimen selection, the evidence strongly supported that humans were the predominate agent of bone accumulation and modification at the site. Furthermore, both bone surface modifications and skeletal elements frequency implied that foragers were most probably engaged in primary butchery of animal carcasses at the site, and subsequently transported selected skeletal elements to a base camp for further processing. The Shiyu site thus can be broadly interpreted as a river-adjacent kill-butchery site. Additionally, we identified a selection of limb bone fragments with distinctive percussive marks as a kind of expedient bone tools, known as bone retouchers. However, regarding the bone fragments from the site previously suspected by some scholars to be bone tools due to their successive scars, it is more reasonable to conclude that these were merely unintended by-products of marrow extraction processes. It is also worth mentioning that a few deliberately produced linear marks were discovered, clearly distinguishable from typical butchery cut marks, undoubtedly reflecting symbolic behaviors and the complexity of human activities to a certain degree.

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    Survival stress of the Yuan Dynasty population in Guangrao, Shandong
    GUO Mingxiao, ZHAN Senyang, ZENG Wen, YOU Haijie, SONG Meiling
    Acta Anthropologica Sinica    2025, 44 (02): 270-282.   DOI: 10.16359/j.1000-3193/AAS.2024.0102
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    This paper presents an archaeological study of human bone remains and stable isotope analysis of 19 Yuan Dynasty human bone specimens excavated from the Shicun North Cemetery in Guangrao, Shandong in 2021. Based on burial items and tomb layout, it is concluded that the Shicun North Cemetery dates back to the Yuan Dynasty. With a minimal quantity of mostly common burial objects per tomb, this cemetery is identified as a civilian cemetery. The research aims to gain insights into the survival stress of the people in Guangrao, Shandong during the Yuan Dynasty.

    Three individuals had fractures, including lumbar vertebra compression fractures, metatarsal fractures, and frontal bone fractures. The frontal bone’s depressed fracture, potentially caused by violence, was relatively rare. This indicates that society was likely stable at that time. There might have been occasional minor disputes, but overall, people lived in peace.

    The results of C and N isotope analysis show that the cemetery population mainly consumed millet and wheat, which is consistent with the grain-consumption structure of the Yuan Dynasty in Shandong, where both millet and wheat were equally emphasized. Meanwhile, the meat supply for the population was relatively abundant and nutritious, yet there were differences in individual consumption levels.

    In the study of survival stress, most people in this cemetery community survived the vulnerable growth and development stages, with only a small number failing to do so due to nutritional status or other factors. To conduct the analysis, the population was divided into adults and minors using osteological paradox and life-history studies. The four minor cases had varying stress-bearing capacities, but none developed strong resistance and adaptability after experiencing severe growth-related stress. One individual was in a vulnerable state and had the strongest pressure-bearing ability among them. Another individual had a weaker pressure-bearing ability compared to others.

    Adult individuals, benefiting from sufficient nutrition, entered a healthy period after reaching adulthood, with high levels of physical and physiological health, strong resilience, and the ability to tolerate and adapt to stress to a certain extent. However, adults also faced higher stress levels because, due to social status and other reasons, they lacked adequate cultural buffering mechanisms.

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    Chronological studies of the Xuetangliangzi hominin fossil site in Hubei Province
    HUA Jiequn, GE Junyi, LU Chengqiu, SHEN Zhongshan, XING Song, LU zeji, GAO Xing, DENG Chenglong
    Acta Anthropologica Sinica    2025, 44 (02): 316-332.   DOI: 10.16359/j.1000-3193/AAS.2025.0015
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    As a key area for the occupation and dispersal of Homo erectus, a large number of Pleistocene H. erectus fossils have been discovered in East Asia. This provides specific advantages for understanding the evolution and dispersal of H. erectus on our planet. However, so far, the scarcity and poor preservation of the early H. erectus fossils ever found in China, especially the lack of completely preserved skull fossils, significantly hinder our knowledge of the evolutionary history of East Asian H. erectus and its evolutionary systematic status.

    Three relatively well-preserved early H. erectus skull fossils have been unearthed from the Xuetangliangzi (Yunxian hominin) site in Hubei province. These fossils hold considerable promise for elucidating this species’ evolutionary trajectory, migratory patterns, and environmental adaptability in East Asia. Consequently, establishing an accurate chronological framework for these fossils is deemed essential.

    Since these skull fossils are of great significance for understanding the evolution, dispersal, and environmental adaptations of East Asian H. erectus, establishing a precise chronological framework for these fossils seems to be vitally important.

    In this study, the previous studies on the stratigraphic and geochronological dating of the Yunxian hominin site were systematically reviewed and examined to analyze the potential problematic issues. At least five levels of terraces are recognizable in the Yunxian hominin site area, and the WT section which was mostly focused on by previous studies may not correspond to the terrace on which the hominin fossils were discovered. Therefore, this discrepancy suggests that many dating efforts, particularly paleomagnetic analyses, may not reliably reflect the site’s antiquity. Electron Spin Resonance (ESR) dating of mammalian remains from the same stratum provides a minimum age estimate for the Yunxian hominin fossils. Furthermore, the incomplete characterization of the mammalian fauna associated with the site complicates faunal comparisons, thereby affecting the chronological assessment.

    Based on this conclusion and integrating regional geomorphological and geochronological data, we propose that disparities in sampling resolution, experimental methodologies, analytical precision, and sample integrity may contribute to the existing controversies. However, the most significant factor undermining the chronological reliability of the Yunxian hominin site is the tenuous nature of regional stratigraphic correlations. Additionally, geochronological datings on the terraces along the Han River indicate that T4 to T1 terraces developed at approximately 55,220,780 and 1,300,000 BP, respectively. The formation of the T4 terrace, where the Yunxian hominin site is located, sets a maximum age constraint for the site. Moreover, a detailed analysis of the temporal distribution of hominin occupations in this region could yield pivotal evidence for correlating the Yunxian hominin fossil with potential cultural affiliations to Paleolithic sites along the river.

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    Reduction patterns and assemblage of the Levallois technology
    WANG Xiaoyu, GAO Xing
    Acta Anthropologica Sinica    2025, 44 (01): 1-13.   DOI: 10.16359/j.1000-3193/AAS.2024.0067
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    Levallois technology, denominated after the Levallois-Perret site located in the northern suburbs of Paris, France, was initially recognized and demarcated by the archaeologist Henri Breuil. This particular technology, which necessitates the fabrication of specific flake products through core preparation, forms a distinctive hallmark of the Middle Paleolithic epoch in the western region of the Old World (referred to as the Middle Stone Age in the African context). It epitomizes the primary technological expression of Mode 3 preformed cores within Clark’s five-mode technological classification system. Marked by the pre-shaping of cores and the generation of specialized flake varieties, Levallois technology held sway in the Mousterian industrial complex.

    Levallois technology originally sprang up during the late phase of the Lower Paleolithic Acheulian industry and subsequently experienced further progression within the Mousterian technological paradigm during the Middle Paleolithic period. The incidence and application of Levallois technology exhibited pronounced variations among diverse Paleolithic assemblages; nevertheless, it unfailingly materialized in a multitude of stone tool industries spanning the Near East, Europe, Africa, and Asia. This lithic technology was noteworthy for its elaborate preforming sequence and the production of standardized implements, attesting to the elevated cognitive capabilities of ancient humans. As a result, this technology is routinely contemplated in the reconstructions of human intellectual evolution, given that it intimates advanced powers of conceptualization, abstraction, intelligence, and language.

    In contradistinction to its widespread prevalence in other regions, the documentary evidence of Levallois technology in China remains comparatively scant, attributable to the paucity of archaeological vestiges. Some scholars previously attributed the deficiency of Levallois technology in East Asia to meager effective population sizes and the absence of the Acheulian heritage. However, in recent years, with the strides made in archaeological excavations, the discovery of artifacts embodying Levallois technology at Chinese sites has galvanized the attention of the academic fraternity. Indicative Levallois artifacts have been unearthed at several locations in northern China, especially in areas proximate to Russia and Mongolia. These sites, dating back approximately 50,000 to 40,000 years ago, include Shuidonggou, Jinsitai, and Tongtian Cave. These findings have subverted prior assumptions and furnished novel vantage points for grasping technological assemblages, technological dissemination, as well as the divergences from representative sites of the late Middle Pleistocene and early Late Pleistocene in northern China.

    This paper strives to introduce and encapsulate the Levallois concepts, products, and idiosyncratic traits of the reduction pattern. It clarifies the extant evidence of Levallois technology unearthed in China and the characteristics of the affiliated industries. By dint of this endeavor, we aspire to proffer valuable perspectives for further research undertakings.

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    Core flaking technology of the Shuijingbei locality of Xiachuan site in Shanxi
    CAI Xinyi, DU Shuisheng
    Acta Anthropologica Sinica    2025, 44 (01): 14-26.   DOI: 10.16359/j.1000-3193/AAS.2024.0049
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    Located in Qinshui County, Jincheng City, Shanxi Province, the Xiachuan site stands out as a significant Upper Paleolithic archaeological site. Excavations have revealed five distinct localities within the site, namely Xiaobaihuageliang, Fuyihegeliang, Liushuiyao, Shuijingbei, and Niulu localities. Two squares exhibiting similar stratigraphic accumulations were excavated at the Shuijingbei locality. The stratigraphic accumulation can be categorized into two primary cultural stages. The lower cultural layer comprises a wealth of remains characterized by a simple core-flake industry, indicative of the early Upper Paleolithic period (44~33 kaBP). The upper cultural stage contains remnants of microblade technology, corresponding to the middle Upper Paleolithic period (26~25 kaBP). During the early Upper Paleolithic stage at the Shuijingbei locality, the prevalent technological approach was simple core-flake industry, most artifacts were directly knapped by hard hammers with a small number of artifacts using bipolar method. In the upper cultural period at the Shuijingbei locality, microblade technology emerged alongside the continued presence of the simple core-flake industry. Among all the excavation localities within the Xiachuan site, the Shuijingbei locality stands out as having the most abundant remains associated with the simple core-flake industry. This particular locality offers valuable insights into the evolutionary trajectory of flaking practices across different time periods. A thorough analysis of the technical attributes of simple core-flake industry cores discovered from these distinct cultural layers at the Shuijingbei locality reveals two distinct flaking organization strategies: Platform orientation and flaking surface orientation. The platform orientation organization encompasses four distinct flaking patterns: Single flaking direction and surface, single flaking direction with multiple flaking surfaces, dual flaking surfaces with a single flaking direction, and multiple flaking surfaces and directions. Conversely, the more sophisticated flaking surface orientation strategy incorporates lateral directions within a single flaking surface and opposing flaking directions within the same flaking surface, constituting two distinctive flaking patterns and highlighting the complexity of flaking techniques. Remnants from both the early and middle Upper Paleolithic periods at the Shuijingbei locality in the Xiachuan site display evidence of these flaking organization strategies, suggesting the enduring nature of the simple core-flake industry’s techniques. Notably, during the middle Upper Paleolithic period at the Shuijingbei locality, advancements in core-flaking techniques are evident, characterized by improved platform retouch techniques, continuous flaking on a single surface, enhanced control over flakes’ shape and size, and increased core development and utilization. By comparing the characteristics of these flaking techniques, it can be inferred that the advancements observed in the middle Upper Paleolithic stage were likely influenced by the concurrent presence of microblade technology during the same period. This interaction between the two flaking technologies reflects the movements and migrations of ancient populations across the vast eastern part of the Eurasian continent, particularly in northern China, during the Upper Paleolithic phase.

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    An experimental archaeological study of the formation causes of incomplete bones from the Wulanmulun site
    TANG Yimeng, LIU Yang, HOU Yamei
    Acta Anthropologica Sinica    2025, 44 (01): 42-54.   DOI: 10.16359/j.1000-3193/AAS.2024.0032
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    Animal skeletons serve as valuable artifacts and crucial research targets in Paleolithic sites. Nevertheless, fragmented animal bones, especially those with species difficult to identify, have yet to receive adequate attention. The Wulanmulun site, a Middle Paleolithic site situated on the banks of the Wulanmulun River in Kangbashi New District, Ordos City, Inner Mongolia Autonomous Region, dates back to 65 kaBP ~ 50 kaBP. Since its discovery and excavation in 2010, a substantial number of stone artifacts and animal fossils have been unearthed. Currently, in-depth investigations have been carried out regarding lithic artifacts, zooarchaeology, and taphonomy at this site. Hence, we will concentrate on the abundant previously unstudied incomplete skeletons and explore their formation processes to uncover their associations with human behavior and natural burial.

    This paper centers on the 57,858 incomplete animal bone fossils excavated from the Wulanmulun site between 2010 and 2014, with the aim of probing into the formative factors of incomplete skeletal remains at the site. By means of quantitative analysis, experimental archaeology, and comparative analysis, we scrutinize the quantity and morphology of incomplete skeletal remains in an endeavor to elucidate the cultural traits and behavioral patterns of ancient humans. The findings suggest that: Firstly, the copious small-sized burned bones were presumably utilized as fuel instead of being the byproducts of roasting meat. Secondly, bone flakes, bone tools, and bone artifacts signify the activities of ancient humans in percussing and retouching bones, which differ from mere smashing for procuring food. Thirdly, through comparative analysis, it is deduced that marrow extraction and bone tool manufacturing coexisted at the Wulanmulun site, and the scarcity of 5 - 10 cm sized incomplete bones is correlated with the bone tool production activities of ancient humans. Fourthly, trampling experiments have verified that the fragmentation of bones caused by human and animal trampling is negligible and does not give rise to a large quantity of incomplete bones.

    Consequently, this study implies that the formation of a large number of incomplete bones at the Wulanmulun site is intimately tied to ancient human activities such as marrow extraction, bone tool manufacturing, and bone burning. The Wulanmulun site comprehensively mirrors the cognitive level and utilization of animal bone resources by ancient humans, who not only harnessed meat resources but also exploited bone resources for marrow consumption, bone tool production, and fuel, exhibiting an efficient resource utilization strategy.

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    Morphological comparison of buccal wear marks on molars between ancient Chinese agricultural and agr-pastoralist populations
    ZHOU Yawei, FU Qingxin
    Acta Anthropologica Sinica    2025, 44 (01): 92-104.   DOI: 10.16359/j.1000-3193/AAS.2024.0068
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    During chewing, teeth rub against phytoliths in food and foreign gravel in the soil, generating diverse dental microwear patterns on the buccal surface of the teeth. The distinct diet structures of populations with different life patterns lead to varying food particles in their diets, which in turn result in different dental microwear. To explore whether differences in the diet structures of populations with different life patterns would manifest as microwear patterns, the author selected the mandibular first molars of 79 individuals from the Wanggou site in Zhengzhou and the Jinmaoyuan site in Datong. The microwear patterns on the buccal side of the teeth were observed using a Leica DVM6M ultra-depth microscope. The results indicated significant differences in microwear patterns between the two groups. For the Wanggou people, whose main economic activity was agriculture, the average number of striations was 36.95, the average length of striations was 236.89 microns, and the average ratio of horizontal to vertical striations (Lh/Lv) was 120.89%. In contrast, for the Jinmaoyuan people under a mixed economy of agriculture and animal husbandry, the average number of striations was 28.26, the average length of striations was 262.55 microns, and the average Lh/Lv ratio was 83.51%. Compared with the Jinmaoyuan people, the Wanggou people exhibited shorter striations, higher density, and a higher Lh/Lv ratio. The average number of striations in the Wanggou people tended to increase with age. There was little disparity in the striation pattern between men and women of the Wanggou people. However, the mean number and average length of striations in the male youth group were smaller than those in the female youth group, suggesting that young men consumed a relatively higher proportion of meat. For the Jinmaoyuan people, the average number of striations and the length of striations gradually decreased with age, indicating that there were fewer hard particles in the diet of the middle-aged group and the physical properties of the food were softer. The Lh/Lv ratio in the prime-age group was notably lower than in other groups, implying that they might consume more meat. The study on the differences in buccal microwear between the two groups revealed no significant difference in the internal microwear patterns between the two populations, indicating a relatively stable diet structure. Nevertheless, the two groups had substantial differences in buccal microwear patterns, demonstrating that different diet structures would induce different micro-attrition patterns. The high proportion of plant food in the diet structure of the Wanggou site inhabitants led to the high density of striations. The diversified subsistence modes and the cold, dry, and sand-prone climatic environment of the Jinmaoyuan site might incorporate more hard exogenous particles in the residents’ food, resulting in longer striations.

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