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    A study overview on tooth wear in ancient populations
    LI Haijun, ZENG Yuxin, ADILIJIANG Waili, NUERMAIMAITI Kadier, ZHANG Hailong, LI Wenying
    Acta Anthropologica Sinica    2025, 44 (02): 333-351.   DOI: 10.16359/j.1000-3193/AAS.2024.0081
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    Tooth wear is closely related to human subsistence activities and is a common physiological phenomenon observed on human remains excavated from archaeological sites. It possesses significant research value in fields such as physical anthropology and archaeology. Currently, numerous scholars both at home and abroad have conducted research on tooth wear in ancient populations, yielding a wealth of findings. However, a systematic review is still lacking.

    This paper reviews and summarizes the common types and influencing factors of tooth wear in ancient inhabitants based on relevant research results from both domestic and international sources. Macroscopically, tooth wear can present morphologically as flat, oblique, rounded, spoon-shaped, cup-shaped, and groove wear. Microscopically, it can be further classified into striations and pits. Factors influencing tooth wear include age, food structure, food processing techniques, socioeconomic types, and specific human behavioral patterns that may lead to distinctive wear phenomena. Moreover, factors such as chewing methods, chewing frequency, occlusal relationships between the upper and lower jaws, and the developmental status of teeth also affect tooth wear to a certain extent.

    Research on tooth wear can be applied to infer the individual age at death, explore the dietary structure and socioeconomic patterns of ancient populations, elucidate ancient labor behaviors and social division of labor, reconstruct the patterns of weaning and feeding in children, as well as the functional restoration of occlusal relationships between the upper and lower dental arches. Based on differences in research purposes and subjects, the academic community has proposed specialized observation methods and grading standards for evaluating the degree of tooth wear. Nevertheless, there is currently no unified grading method or standard for tooth wear.

    Overall, exploring the dietary conditions and subsistence patterns of ancient populations from different archaeological sites in China through tooth wear is a major research focus of ancient human tooth wear studies in China. Significant research achievements have revealed the dietary conditions and subsistence patterns of populations from various archaeological sites. In terms of time, these sites are mainly concentrated from the Neolithic Age to the early Iron Age. Geographically, research on tooth wear in ancient Chinese populations is mainly distributed in the Central Plains and the northwestern border regions of China, with relatively fewer studies in the southern regions. Compared with international research, domestic studies on ancient human tooth wear still face limitations such as narrow research perspectives, lack diversity in research focus and homogeneous content. Future research could benefit from innovations in research fields, ideas, and methods to fill the gaps and achieve further progress.

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    Hominin and human dispersals in palaeolithic East Asia
    Robin DENNELL
    Acta Anthropologica Sinica    2025, 44 (01): 132-164.   DOI: 10.16359/j.1000-3193/AAS.2022.0044
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    Dispersals, colonisation, immigration and population assimilation or replacement are fundamental themes in the Palaeolithic record of East Asia. Some of these issues can be studied within a biogeographic framework that explains why and how the distribution of hominin species changed over time and space in response to climatic and environmental change. Because hominins (and especially humans) can change their behaviour through technical, social and cognitive developments, biogeographic models also have to incorporate this factor when investigating dispersals. This is particularly important with the dispersals in East Asia by Homo sapiens into rainforests, across open sea to off-shore islands, to the Arctic and the highest parts of the Tibetan Plateau. This paper suggests how hominin and human dispersals in East Asia might be investigated by using a biogeographic framework that can incorporate changes in hominin adaptability and behaviour.

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    Reduction patterns and assemblage of the Levallois technology
    WANG Xiaoyu, GAO Xing
    Acta Anthropologica Sinica    2025, 44 (01): 1-13.   DOI: 10.16359/j.1000-3193/AAS.2024.0067
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    Levallois technology, denominated after the Levallois-Perret site located in the northern suburbs of Paris, France, was initially recognized and demarcated by the archaeologist Henri Breuil. This particular technology, which necessitates the fabrication of specific flake products through core preparation, forms a distinctive hallmark of the Middle Paleolithic epoch in the western region of the Old World (referred to as the Middle Stone Age in the African context). It epitomizes the primary technological expression of Mode 3 preformed cores within Clark’s five-mode technological classification system. Marked by the pre-shaping of cores and the generation of specialized flake varieties, Levallois technology held sway in the Mousterian industrial complex.

    Levallois technology originally sprang up during the late phase of the Lower Paleolithic Acheulian industry and subsequently experienced further progression within the Mousterian technological paradigm during the Middle Paleolithic period. The incidence and application of Levallois technology exhibited pronounced variations among diverse Paleolithic assemblages; nevertheless, it unfailingly materialized in a multitude of stone tool industries spanning the Near East, Europe, Africa, and Asia. This lithic technology was noteworthy for its elaborate preforming sequence and the production of standardized implements, attesting to the elevated cognitive capabilities of ancient humans. As a result, this technology is routinely contemplated in the reconstructions of human intellectual evolution, given that it intimates advanced powers of conceptualization, abstraction, intelligence, and language.

    In contradistinction to its widespread prevalence in other regions, the documentary evidence of Levallois technology in China remains comparatively scant, attributable to the paucity of archaeological vestiges. Some scholars previously attributed the deficiency of Levallois technology in East Asia to meager effective population sizes and the absence of the Acheulian heritage. However, in recent years, with the strides made in archaeological excavations, the discovery of artifacts embodying Levallois technology at Chinese sites has galvanized the attention of the academic fraternity. Indicative Levallois artifacts have been unearthed at several locations in northern China, especially in areas proximate to Russia and Mongolia. These sites, dating back approximately 50,000 to 40,000 years ago, include Shuidonggou, Jinsitai, and Tongtian Cave. These findings have subverted prior assumptions and furnished novel vantage points for grasping technological assemblages, technological dissemination, as well as the divergences from representative sites of the late Middle Pleistocene and early Late Pleistocene in northern China.

    This paper strives to introduce and encapsulate the Levallois concepts, products, and idiosyncratic traits of the reduction pattern. It clarifies the extant evidence of Levallois technology unearthed in China and the characteristics of the affiliated industries. By dint of this endeavor, we aspire to proffer valuable perspectives for further research undertakings.

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    An experimental study of human tooth marks on bones
    CAO Yuxin, SUN Lu, ZHANG Yue, ZHANG Shuangquan
    Acta Anthropologica Sinica    2025, 44 (02): 242-254.   DOI: 10.16359/j.1000-3193/AAS.2024.0053
    Abstract646)   HTML56)    PDF(pc) (4250KB)(658)       Save

    In past decades, researchers primarily focused on the distinguishing characteristics of marks on bone surfaces, such as cut marks, percussion marks and categories of modifications, which are widely considered as direct indicators ancient human behaviors. While earlier investigations often attributed tooth markings found on animal remains to carnivore or rodent consumption habits, recent work by Western archaeologists and ethnoarchaeological researchers consistently demonstrates that humans can inflict a category of modifications on bones during their chewing. In such studies, researchers have also attempted to find ways to distinguish human tooth marks from those produced by other carnivores through actualistic experiments. However, few scholars in China have focused on identifying human tooth marks on animal remains. The current study is thus serving as a complementary work upon prior investigations and at the same time adding to the growing evidence to show that humans do indeed leave discernible dental imprints upon animal skeletal remains during consumption processes—particularly upon flatter or more fragile bones. Damages observed on bones include furrowing, scooping-out, crenulated and saw-toothed edges, longitudinal cracking, crushing, peeling and score. In this paper, we present a detailed description of the types of damages observable on bone surfaces in an experimental human chewing of the bones of sheep. A total of 167 chewed post-cranial bones from different parts of sheep have been examined, including 122 boiled bones and 45 roasted ones. Our experimental observation indicates that, although there is a certain degree of morphological similarity between human tooth marks and those marks caused by carnivores, the former assemblage could still be confidently identified and it thus has the potential to provide us with an independent standard for identifying human actors in faunal remains from the archaeological hominins. On the other hand, the description of human chewing patterns may provide further evidence of exploitation of certain groups of animals where cut-marks are rare, such as small games or birds. Furthermore, this experiment provides valuable materials for the comparative analysis and of human tooth-marked from the Chinese archaeological sites. Additionally, the types of tooth marks observed in this experiment are largely consistent with the findings of Western scientists, indicating that these observations have a considerable degree of universality and can be applied to the study of archaeological materials across a broader geographical range. The present paper is the first attempt made by Chinese archaeologists to document human tooth marks in an actualistic setting and it has the potential to provide us with a novel approach to explore human subsistence strategies in Paleolithic China.

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    Exploration of "behavioral modernity” and “modern human behavior” in human evolution
    YANG Shixia
    Acta Anthropologica Sinica    2025, 44 (02): 181-192.   DOI: 10.16359/j.1000-3193/AAS.2025.0014
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    The evolution and widespread of Homo sapiens is a key event in the human evolution history. By the end of the Late Pleistocene, our species was distributed across every continent except Antarctica, and adopted to varied ecological niches. The behavioral complexity and the cultural diversity are believed to act as one of the most pronounced abilities to get success. The fossil and genetic evidence were applied to discuss the extinction of archaic populations, such as the Denisovans and the Neanderthals and their complicated interbreeding with Homo sapiens. The archaeological records served on the discussion about the behavioral evolution from the emergence of “behavioral complexity or modernity’ to the fully developed modern human ‘behavioral package’. In this paper, we reviewed the discussion on the modern human behavioral evolution in the western side of the Old World. Based on the African archaeological records, the behavioral origins of Homo sapiens can be traced back to the first material culture produced by our species in Africa and phased as Middle Stone Age (MSA); while the western Eurasian records the behavioral complexity raised with the origin of Neanderthals and phased as Middle Paleolithic (MP). Then around the middle of Late Pleistocene, as the wide distribution of Homo sapiens and the extinction of archaic populations, a complete ‘modernity package’ finally developed and shows more cultural diversity globally, which including the advanced lithic tools (blade, microblade), bone tools, increased geographic range, specialized hunting, the use of aquatic resources, long distance exchange, systematic processing and use of pigment, and art and decoration.

    However, recently in Eastern Asia efforts were made to explore the complexity of the ‘behavioral complexity or modernity’, the technological innovations and behavioral adaptations process are still barely known. The lithic assemblages have long been regarded as simple, conservative technologies, and major changes in raw material procurement, core reduction, retouch and typology tended to be clustered until 40 kaBP, without a ‘middle phase’ showing distinguished transition. These years, facing with the high degree of hominin morphological variability which has been suggested by recent fossil and genetic evidence, such as the early modern humans, Denisovans, H. longi, and some other unassigned taxons (Xujiayao and Xuchang), and growing new archeological data, we need to rethink the ‘behavioral modernity’ process happened in Eastern Aisa and its application to the evolutionary history of our species worldwide.

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    Progress and perspectives of the isotope research of human tooth enamel
    LEI Shuai
    Acta Anthropologica Sinica    2025, 44 (01): 165-180.   DOI: 10.16359/j.1000-3193/AAS.2024.0019
    Abstract589)   HTML49)    PDF(pc) (1829KB)(588)       Save

    This paper meticulously addresses the significant theoretical and methodological aspects involved in isotopic investigations of human tooth enamel. Firstly, it systematically organizes the growth and development mechanism, principal chemical composition, and the efficacy of human tooth enamel in withstanding diagenesis. Emphasis is placed on the inherent advantages that have rendered it the primary target for hydroxyapatite isotope analyses and studies within the current bioarchaeological realm. Secondly, a vast array of research topics centered around the multi-isotope analysis of elements such as oxygen (δ18O), carbon (δ13C), strontium (87Sr/86Sr), zinc (δ66Zn), magnesium (δ26Mg), calcium (δ44/42Ca), and lead (Pb) in human enamel are distilled and consolidated into three scientific themes: human life history, habitation, and health. Additionally, the current status of each of these research areas is comprehensively reviewed. Finally, within the framework of constructing the theoretical system of archaeology with Chinese characteristics in the new era, the exigency of the swift advancement of multi-isotope research on tooth enamel is underscored. In light of the existing research experiences and deficiencies in the international bioarchaeology community, future research in China is anticipated to progress from the vantage points of innovating experimental sampling strategies and research methodologies, accurately discerning the merits and demerits of diverse mass spectrometry techniques, broadening the depth and scope of research topics, and delving deeply into the cooperative interactions between isotopes and different human body tissues.

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    Hunting and utilization of roe deer by prehistoric humans at the Honghe site in Heilongjiang
    LIANG Qiyao, YUE Jianping, ZHANG Wei, CHEN Quanjia
    Acta Anthropologica Sinica    2025, 44 (01): 105-116.   DOI: 10.16359/j.1000-3193/AAS.2024.0101
    Abstract521)   HTML52)    PDF(pc) (2289KB)(555)       Save

    The Honghe site (123°35′56″E, 47°06′49″N), situated on the right bank of the middle reaches of the Nenjiang River in Qiqihar, Heilongjiang Province, Northeast China, has been ascertained to be a pivotal location for comprehending prehistoric and historical human activities in this region. Between 2013 and 2019, the Heilongjiang Provincial Institute of Cultural Relics and Archaeology carried out five excavations at the Honghe site, leading to the discovery of copious cultural relics and faunal remains. These findings span an extensive chronological range, from the Late Neolithic to the Qing Dynasty, underlining the protracted history of human settlement and resource exploitation in the area. Among the diverse animal remains unearthed, the bones of roe deer (Capreolus pygargus) stand out notably.

    The principal aim of this study was to analyze the roe deer bones excavated from the Honghe site, with the intention of discerning the interactions between humans and roe deer during the Late Neolithic era. By conducting a comprehensive examination of the roe deer skeletal remains, researchers endeavored to reconstruct the modalities of hunting, processing, and utilization by ancient inhabitants. The analysis incorporated the investigation of the age structure at death, sex ratios, seasonal hunting patterns, skeletal distributions, and techniques of bone processing and utilization. The results demonstrated that the mortality of roe deer predominantly transpired within two age brackets: Prime ages of 3~7 years and 7~9 years, and a higher proportion of males was detected among the specimens. This suggests that hunters might have deliberately targeted these age groups, potentially due to their size and the quality of their antlers.

    Hunting activities at the Honghe site seemingly took place throughout the year, indicating a continuous dependence on roe deer as a resource. Although seasonal fluctuations were observed, the data imply that roe deer served as a dependable source of both meat and bone materials year-round. The bones most commonly unearthed comprised mandibles, antlers, and limb bones, affording glimpses into the portions of the animal that were most prized. The presence of antlers intimates that hunters may have focused on males for their antlers, which could have been employed for fabricating tools and ornaments.

    Furthermore, the site yielded an assortment of bone tools fashioned from roe deer bones, such as fishhooks, awls, and daggers. These artifacts accentuate the role of roe deer bones as a crucial raw material for tool manufacture, exemplifying the adaptability and ingenuity of the Neolithic inhabitants. It is highly probable that the roe deer hides played a significant part in the production of clothing, furnishing warmth and safeguarding against the frigid climate of Northeast China.

    In toto, the study of the Honghe site illuminates the significant part that roe deer played in the subsistence and material culture of the ancient inhabitants of this region. The findings offer precious insights into the broader fishing and hunting economy of the Late Neolithic period in Northeast China, stressing the importance of roe deer as a multifaceted resource. Through this analysis, researchers have attained a more profound understanding of the means by which the Neolithic communities acclimated to and thrived in their environment, relying on local fauna to support their livelihoods.

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    Ancient DNA reveals the utilization of wild animal resources by prehistoric humans in Northwest China
    SONG Guangjie, CAI Dawei, ZHU Cunshi, HU Songmei, ZHOU Jing, REN Xiaoyan
    Acta Anthropologica Sinica    2025, 44 (01): 117-131.   DOI: 10.16359/j.1000-3193/AAS.2024.0031
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    Genomic analysis of ancient wild animal remains is of not only great significance for the conservation and utilization of genetic resources of wild animal species but also crucial for helping us understand the diet compositions and hunting activities of ancient human beings as well as their social-economic development patterns. Ancient DNA technology has been widely employed in archaeological research. Among its numerous strengths, its potential to decipher the genetic information carried by biological samples at the molecular level has been widely acknowledged, and many researchers have utilized ancient DNA analysis to distinguish between domestic and wild animals. Moreover, when combined with historical and archaeological evidence, it offers us robust scientific and technological support, enabling us to comprehensively understand ancient human societies, including their origins and evolutionary processes.

    In this study, the ancient DNA of nine animal samples, which were excavated from the Changning, Mogou, Quanhucun, and Dashigou sites in Northwest China and morphologically identified as either “sheep” or “goats”, was investigated using ancient DNA technology. Ancient DNA extraction, library construction, and high-throughput sequencing were carried out, and the mitochondrial genome sequences of the nine samples were successfully obtained. Alignment analysis was performed between the genomic sequences of these samples and the 146 mitochondrial genomic sequences of Cervidae and Bovidae (used as reference data). The results of the alignment analysis indicated that these nine samples were identified as belonging to four different wild animal species within the families of Cervidae and Bovidae: the Siberian roe deer (Capreolus pygargus) of the genus Capreolus within the subfamily Odocoileinae; the Przewalski’s gazelle (Procapra przewalskii) and Tibetan gazelle (Procapra picticaudata) of the genus Procapra within the subfamily Antilopinae; and the Sumatran serow (Capricornis sumatraensis) of the genus Capricornis within the subfamily Caprinae.

    Two phylogenetic analyses were conducted on the mitochondrial genome data extracted from the Odocoileinae, Caprinae, and Antilopinae subfamilies. Bayesian phylogenetic trees and Maximum Likelihood phylogenetic trees were constructed. It was demonstrated that each of the nine samples clustered with the corresponding species identified by the alignment analysis, which was consistent with the results of the principal component analysis on the same data set, where the nine samples were assigned to the corresponding species identified in the alignment analysis. Genetic distance calculations between individuals based on ancient and modern samples revealed that each of the nine samples was genetically closest to the specific species identified.

    All of the above results emphasize that ancient DNA technology can overcome the limitations of morphological methods in the species identification of ancient animals. Considering other wild animal bone remains excavated from the four sites, it can be concluded that the ancient people in prehistoric Northwest China used wild animals as a supplement to domestic animal resources for food, sacrificial offerings, and bone tool manufacture. This study is of great significance as it provides new evidence at the molecular level and corroborates the findings of previous archaeological research on animals in prehistoric times.

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    Paleolithics excavated from Area B of the Yahuai Cave site, Guangxi
    LI Hao, XIE Guangmao, LI Dawei, LI Sanling, XIAO Peiyuan, CHEN Qingyi, WANG Yuqing, JIA Zhenxiu, WU Yan, LU Jieying
    Acta Anthropologica Sinica    2025, 44 (03): 404-412.   DOI: 10.16359/j.1000-3193/AAS.2025.0047
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    The Yahuai Cave site is a newly discovered Late Paleolithic to Neolithic site in South China. The site has yielded rich cultural remains spanning a relatively long period. Three Paleolithic cultural phases have been identified from the site. Analysis of lithic assemblages shows that two distinctive flaking systems co-existed at the site. One is featured by the steep-angled flaking cores made on sandstone, while the other is featured by the bipolar flaking cores made on tektite, chert, quartz and crystal. Bipolar technology is mainly related to the production of miniaturized flakes and flake tools. Miniaturized lithic assemblages or minialithic technology have been widely documented in the Upper Paleolithic sites in South and Southeast Asia, as well as the different regions of China, likely indicating the migration and dispersal process of early modern humans. The study of knapped stone artifacts at Yahuai Cave provides important materials and evidence for understanding the behavioral diversity and complexity of early modern humans in East Asia.

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    Craniofacial characteristics of She people in Fujian
    HU Rong, YIN Jiahao
    Acta Anthropologica Sinica    2025, 44 (04): 631-641.   DOI: 10.16359/j.1000-3193/AAS.2025.0005
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    This study randomly selected 518 She ethnic adults aged 18 years and above (293 males and 225 females) from Fu’an and Fuding cities in Fujian Province. Twenty-two craniofacial characteristics were measured, and twelve indexes of craniofacial exponent characteristics were calculated. The results showed that the She ethnic group has a medium head type, high head type, and narrow head type; a narrow face; and a medium nose type. There were significant gender differences in the craniofacial characteristics and indexes of the She ethnic group in Fujian, except for the intraocular angular width and zygomatic frontal index. There were significant age differences in head width, face width, intraocular width, lip - related indicators (upper lip height, lip height, lip thickness, mouth breadth), ear length, vertical cephalo - facial index, and oral index. Facial features are mainly determined by genetic factors.
    When comparing the craniofacial characteristics of the She ethnic group in Fujian with those of the She ethnic group 28 years ago, the results showed that the craniofacial characteristics of the She ethnic group in this study had changed significantly compared to those 28 years ago. The reasons were related to urban - rural differences, environmental and dietary changes, and an increase in intermarriage with the Han and other ethnic groups.
    Principal component analysis and cluster analysis were conducted on the head - face data of the She, Han, and other ethnic minority populations. It was found that the She people, as an independent group, were between the northern ethnic minorities, southern ethnic minorities, and the Han. Furthermore, the craniofacial characteristics of the She in Fujian were closer to those of the Han in South China, the Miao, the Yao, the Shui, and the Wa, and had significant differences from other southern ethnic minorities in Yunnan, Guangxi, Guizhou, and northern ethnic groups.
    The She ethnic group has long lived in the border areas of Guangdong, Fujian, and Jiangxi in South China. Therefore, compared to the Han and minority ethnic groups in North China, the Han and minority ethnic groups in South China have more similar physical characteristics to the She ethnic group. The migration and integration of ethnic groups is an important historical process in the formation of ethnic groups in history. The genetic and cultural blending between ethnic groups has influenced their physical characteristics. This paper, together with evidence from Ethnology, Molecular Anthropology, and Linguistics, shows that the She ethnic group had intricate connections with minority ethnic groups such as the Miao - Yao, Min, Yue, as well as the Han ethnic group in the past. People with different physical characteristics participated in this process and ultimately formed the modern She ethnic group’s “diverse and integrated” physical characteristics.

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    Morphological comparison of buccal wear marks on molars between ancient Chinese agricultural and agr-pastoralist populations
    ZHOU Yawei, FU Qingxin
    Acta Anthropologica Sinica    2025, 44 (01): 92-104.   DOI: 10.16359/j.1000-3193/AAS.2024.0068
    Abstract463)   HTML37)    PDF(pc) (1961KB)(396)       Save

    During chewing, teeth rub against phytoliths in food and foreign gravel in the soil, generating diverse dental microwear patterns on the buccal surface of the teeth. The distinct diet structures of populations with different life patterns lead to varying food particles in their diets, which in turn result in different dental microwear. To explore whether differences in the diet structures of populations with different life patterns would manifest as microwear patterns, the author selected the mandibular first molars of 79 individuals from the Wanggou site in Zhengzhou and the Jinmaoyuan site in Datong. The microwear patterns on the buccal side of the teeth were observed using a Leica DVM6M ultra-depth microscope. The results indicated significant differences in microwear patterns between the two groups. For the Wanggou people, whose main economic activity was agriculture, the average number of striations was 36.95, the average length of striations was 236.89 microns, and the average ratio of horizontal to vertical striations (Lh/Lv) was 120.89%. In contrast, for the Jinmaoyuan people under a mixed economy of agriculture and animal husbandry, the average number of striations was 28.26, the average length of striations was 262.55 microns, and the average Lh/Lv ratio was 83.51%. Compared with the Jinmaoyuan people, the Wanggou people exhibited shorter striations, higher density, and a higher Lh/Lv ratio. The average number of striations in the Wanggou people tended to increase with age. There was little disparity in the striation pattern between men and women of the Wanggou people. However, the mean number and average length of striations in the male youth group were smaller than those in the female youth group, suggesting that young men consumed a relatively higher proportion of meat. For the Jinmaoyuan people, the average number of striations and the length of striations gradually decreased with age, indicating that there were fewer hard particles in the diet of the middle-aged group and the physical properties of the food were softer. The Lh/Lv ratio in the prime-age group was notably lower than in other groups, implying that they might consume more meat. The study on the differences in buccal microwear between the two groups revealed no significant difference in the internal microwear patterns between the two populations, indicating a relatively stable diet structure. Nevertheless, the two groups had substantial differences in buccal microwear patterns, demonstrating that different diet structures would induce different micro-attrition patterns. The high proportion of plant food in the diet structure of the Wanggou site inhabitants led to the high density of striations. The diversified subsistence modes and the cold, dry, and sand-prone climatic environment of the Jinmaoyuan site might incorporate more hard exogenous particles in the residents’ food, resulting in longer striations.

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    Perforated stone tools from the Yahuai Cave site in Long’an, Guangxi
    XIE Ying, FU Lin
    Acta Anthropologica Sinica    2025, 44 (03): 439-450.   DOI: 10.16359/j.1000-3193/AAS.2025.0036
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    Perforated stones are widely distributed in prehistoric sites across South China and Southeast Asia. Typically coexisting with chipped stone tools, ground stone tools, and other relics, they constitute a significant element of the regional prehistoric culture. However, due to the limited number of perforated stones unearthed at each site, in-depth research opportunities are substantially restricted, leading to diverse academic viewpoints regarding their functions and usage.

    In Guangxi, the perforated stones excavated date from the late Paleolithic to the late Neolithic period, indicating their long - term utilization. In recent years, a substantial quantity of perforated stones has been discovered at Yahuai Cave in Guangxi. These artifacts not only date back to as early as 25,000 years ago but also exhibit diverse shapes and distinct usage traces.

    This study conducts a comprehensive analysis of the shape characteristics, raw materials, and usage traces of the perforated stones from Yahuai Cave. The size characteristics of the perforated stones and the presence or absence of a grinding surface serve as the classification criteria. Archaeological evidence at the site reveals that prehistoric inhabitants of Yahuai Cave utilized stone anvils to process these perforated stones. The raw materials of the Yahuai Cave perforated stones are predominantly flat, round, or nearly round sandstone cobbles. They were perforated through a two - face piercing method at the center of the stone body, with stone awls employed as the primary processing tool.

    Most of the perforated stones display usage traces in the hole area and on parts of the stone body, and some are stained with ochre pigment. Integrating the site context, this study reveals the functional diversity of these perforated stones. It is speculated that they might have been used for plant rope making, as fire - making tools, and digging tools. Additionally, they could have served as stone hammers, stone anvils, and stone grinding tools, participating in ochre processing activities such as grinding and crushing. The extensive usage traces on various parts of the perforated stones suggest a high overall utilization rate, indicating that they were versatile utility tools.

    On one hand, the functional diversity of perforated stones reflects the innovative tool - using practices of prehistoric people; on the other hand, it also demonstrates the wisdom and environmental adaptation strategies of prehistoric communities.

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    Research on the tektite artifacts unearthed at the Yahuai Cave site in Guangxi
    GUO Yunhao, CHEN Hong, LI Dawei, XIE Guangmao, XUE Liping
    Acta Anthropologica Sinica    2025, 44 (03): 451-465.   DOI: 10.16359/j.1000-3193/AAS.2025.0049
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    Tektites are frequently discovered in prehistoric sites in Guangxi, yet research on the associated artifacts remains limited. This study conducts an in-depth archaeological analysis of the tektite artifacts unearthed at the Yahuai Cave site in Guangxi, exploring their role and significance in the lives of ancient humans. These tektite specimens originate from the Australasian strewn field and exhibit identical provenance to those tektites recovered from sedimentary strata within Baise Basin, Guangxi. Their formation correlates with the Australasian tektite event, dating to approximately 700,000~800,000 years before present. A comprehensive approach we employed that includes methods such as technological analysis, statistical analysis, energy dispersive X-ray spectroscopy (EDXRF) analysis,and use-wear analysis to investigate the physical and chemical characteristics, flaking strategies, retouch logic, post-depositional modification, and usage conditions of the tektite assemblages.

    The results of EDXRF analysis demenstrate that the tektites from the Yahuai Cave site are chemically consistent with those from surrounding areas. Regarding to the tool manufacturing, two flaking strategies were employed: the bipolar knapping and the "strike-rotate" knapping. Additionally, comprehensive and meticulous or localized retouched methods were adopted based on actual needs, demonstrating a profound understanding and adaptation to the material properties. It also reveals the optimization and improvement in stone tool manufacturing by ancient people. The evaluation of post-depositional modification indicates that the tektites experienced a certain degree of wear during the post-depositional process, but it did not significantly affect the use-wear traces of their surfaces.

    The analysis of usage logic suggests that the use of tektites at the Yahuai Cave site was primarily related to the development and exploitation of animal resources, possibly closely associated with hunting and butchering activities. These findings not only provide new insights into the trend of miniaturization of stone artifacts in southern regions but also offer valuable information for understanding how ancient humans adapted to environmental changes, utilized resources, and the evolution of social structures.

    Overall, this article discusses the uniqueness of these tektites and their prevalence in the Yahuai Cave site, highlighting their important role in ancient toolkits. The results suggests that the utilization of tektites reflects the ability of ancient humans to adapt to their environment and make economic strategy choices. Furthermore, it provides a new perspective and evidence for exploring both the trend of stone artifacts miniaturization in southern China and the dispersal routes of modern humans.

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    An excavation report of the Yahuai Cave site in Long’an, Guangxi
    XIE Guangmao, LIN Qiang, YU Minghui, LU Jieying
    Acta Anthropologica Sinica    2025, 44 (03): 365-388.   DOI: 10.16359/j.1000-3193/AAS.2025.0046
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    The Yahuai Cave site is situated at a limestone tower near Bolang Village, Long’an County, Guangxi Zhuang Autonomous Region, South China (107°45′35″E, 23°6′25″N). It is a prehistoric site predominantly characterized by Palaeolithic deposits. Between 2015 and 2018, the Guangxi Institute of Cultural Relic Protection and Archaeology carried out excavations at this site. Four areas, namely A, B, C, and D, were opened, with a total exposure area of 50 m². A burial containing a complete Palaeolithic human skull was discovered. A multitude of cultural remains, such as chipped stone tools, ground stone tools, shell and bone artifacts, and potsherds, were unearthed. The majority of the stone tools are made of small quartz sandstone, tektite, chert, or crystal stones, exhibiting features typical of the “core-flake industry” commonly found in North China. Three hearths and an ash pit were identified, suggesting frequent fire usage at the site. Ochres were found throughout all cultural layers, and some of them were discovered to be ground into powder. Numerous bones of terrestrial and aquatic animals were excavated. The site was dated using the AMS 14C dating method to a period between 43,000 and 4,000 BP cal. Four cultural phases were recognized, with phases 1~3 belonging to the Palaeolithic period and phase 4 to the Neolithic period.

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    Paleopathology of human bones from the pottery site of Dayuan Village in Shaanxi
    CHEN Liang, SONG Yuke, FU Zhongyang, XU Lianggao
    Acta Anthropologica Sinica    2025, 44 (01): 55-65.   DOI: 10.16359/j.1000-3193/AAS.2024.0055
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    This paper undertakes Physical Anthropological research on the excellently preserved skeletal remains of 30 cases unearthed from the pottery site of the Western Zhou Dynasty in Dayuan Village, Xixian New Area, Xi’an City, Shaanxi Province. Through a combination of naked-eye observation and Dino-Lite microscopy, the paleopathological manifestations are meticulously recorded. The pottery site in Dayuan Village represents a significant archaeological discovery in the Xi’an region, predominantly yielding sand-tempered brown pottery “split crotch” li tripods. The products fabricated at this site are relatively homogeneous and display a high degree of specialization.

    The pathological findings disclose that cribra orbitalia was prevalent among the majority of the ancient inhabitants, with some cases accompanied by porotic hyperostosis. This suggests that the local population might have endured iron-deficiency anemia, suffered from malnutrition, and resided in a poor living environment. A high incidence of rib periostitis and maxillary sinusitis was observed among the ancient site residents, potentially attributable to respiratory ailments induced by environmental pollution. The ancient inhabitants of this pottery site exhibited a remarkable frequency of metatarsal-phalangeal joint osteoarthritis, signifying the frequent utilization of metatarsophalangeal joints. Osteoarthritis, being the most common joint disorder, predominantly affected the spine joints, ankle joints, shoulder joints, and hip joints, reflecting a substantial labor burden. The metacarpals of the ancient residents were anomalously curved, and skeletal evidence of Musculoskeletal Stress Markers was detected on the phalanges and limb bones. These distinctive bony alterations imply that these individuals were involved in repetitive joint motions and occupations that placed significant demands on hand musculature.

    Based on the paleopathology of the population at the site, the following deductions are made. In light of the pathological signs of the ancient residents in the pottery site in Dayuan Village, a high prevalence of cribra orbitalia, porotic hyperostosis, rib periostitis, and maxillary sinusitis was noted within the site population. The prevalence of osteoarthritis and the occurrence rate of metatarsal-phalangeal joint osteoarthritis were also considerable. It is hypothesized that these pathological changes might be associated with the joint stress resulting from long-term strenuous labor and the adverse environment of the pottery site. Considering the overall circumstances, these disease characteristics are intimately linked to the occupation of pottery making. The ancient residents who once resided in Dayuan Village were presumably pottery artisans and engaged in handicrafts over an extended period.

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    Sex and age-at-death of the Yahuai Cave Man from the Late Pleistocene of Guangxi
    HE Letian, XIE Guangmao, LIN Qiang, LI Dawei, WU Xiujie
    Acta Anthropologica Sinica    2025, 44 (03): 389-403.   DOI: 10.16359/j.1000-3193/AAS.2025.0034
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    Obtaining accurate information about an individual’s sex and age at death constitutes the cornerstone of biological anthropological research. The estimation of adult sex from skeletal samples relies on two types of characteristics: those associated with functional differences between the sexes, and those manifested as variations in the size and shape of bones and teeth. Most methods for age estimation are based on the assessment of osteological degenerative changes. The identification of sex and age in human fossils often sparks controversy due to their evolutionary traits and preservation limitations, particularly when the hip bone is unavailable.

    This study utilized visual observation, measurement, and micro-CT scanning to examine the Late Pleistocene crania, mandible, and teeth (YH1) excavated from the Yahuai Cave site in Guangxi, southern China. Regarding non-metric characteristics, the cranial and mandibular morphology indicates that Yahuai 1 is more likely to be female. However, certain robusticity-related features are more pronounced in YH1 compared to most recent-modern females in East Asia. The sexual dimorphism observed in late Pleistocene human skulls differs from that of modern humans, as evidenced by cranio-facial metric features. Comparisons with late Pleistocene human skulls from East and Southeast Asia reveal that YH1 shows no obvious sexual dimorphism in craniometric characteristics. When compared with recent populations from the same region, YH1 is found to be larger in size, and exhibits similarity to recent female samples after size calibration. YH1 has a medium-sized mandible overall and a high corpus robusticity index, which is comparable to that of late Pleistocene and Neolithic females.

    In terms of age-at-death estimation, YH1 displayed a low degree of synostosis in the cranial sutures, and no age-associated features were observed. Moreover, by comparing with two Neolithic populations from South China, the relationship between dental wear and age estimation was adjusted. Collectively, these findings suggest that YH1 was not elderly, but rather a young adult aged between 30 and 40 years.

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    A study of the miniaturized stone artifacts from the Yahuai Cave site in Long’an, Guangxi
    DENG Wanwen, XIE Ying
    Acta Anthropologica Sinica    2025, 44 (03): 413-426.   DOI: 10.16359/j.1000-3193/AAS.2025.0040
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    The Yahuai Cave site in Long’an, Guangxi, preserves stratified deposits dated to 43~14 kaBP cal, yielding a significant collection of lithic artifacts predominantly reflecting a simple core-flake reduction strategy. The lithic raw materials include quartz sandstone, sandstone, tektite, quartz, and flint. Hard-hammer percussion and bipolar techniques were primarily employed for flake production, with cores predominantly exhibiting single-platform, unidirectional reduction. Some cores show minimal preparation of platform edges prior to flaking. Retouched tools, primarily on flake blanks, were modified via unifacial hard-hammer percussion, with small portions of marginal edge retouch. The tool kit is dominated by scrapers (mostly <5 cm in size), alongside choppers and points, collectively representing a miniaturized lithic industry characteristic of the Late Paleolithic in the Lingnan region.

    Similar miniaturized lithic industries have been documented at contemporaneous sites in Lingnan, the middle-lower reaches of the Yangtze River, the Yunnan-Guizhou Plateau, and Southeast Asia. Comparative analyses reveal that these small flake tool industries in Yahuai Cave and adjacent areas predominantly utilized flint, supplemented by lower-quality quartz, crystal, and other fine-grained raw materials. The preferential selection of flint and other fine-grained raw materials marks a distinctive feature of late Late Pleistocene small flake tool industries in South China. Notably, certain sites, such as the Bailian Cave in Liuzhou, Xianrendong and Diaotonghuan Cave sites in Jiangxi, and some others in the Lishui River valley of Hunan, sourced flint from distant outcrops up to 20 km away. In contrast, other sites in Guizhou, including the Da’erwo rockshelter site in Kaiyang, Laoyadong Cave site in Bijie, and Qingshuiyuan Dadong Cave site, utilized locally available materials such as flint, crystal, basalt, and siliceous limestone from nearby bedrock sources. It should also be noted that the early deposits of Yahuai Cave contain both small flint tools and retouched flakes detached from quartz sandstone cobbles. The use of small cobble flakes was also observed at the Nguom rockshelter site in northern Vietnam.

    Technologically, small flake industries across Lingnan and neighbouring regions during the late Late Pleistocene are characterized by hard-hammer direct percussion, unidirectional retouch, and short, shallow, and overlapping retouch scars. Some micro-sized lithics may reflect bipolar reduction, with no evidence of prepared core technologies. At the Yahuai Cave site, cobble cores with steep edge angles exhibit unidirectional flake removal surfaces resulting from intensive, hierarchical reduction. The shared types of raw materials and technological patterns, combined with the small dimensions of retouched tools, correspond to the global trend of lithic miniaturization during the Late Pleistocene.

    Although these miniaturized lithic artefacts in South China and Southeast Asia represent simple core-flake technologies, they may have functioned as composite tool components, as well demonstrated by examples from the Ille Cave site in northern Palawan, Philippines. Moreover, the Xiamabei site in the Nihewan Basin (North China) provides evidence of composite tool use associated with early modern human behavioural complexity. The presence of ochre at the Yahuai Cave site further supports the emergence of complex cultural behaviours.

    Scholars have debated the origins of these small flake industries in South China-Southeast Asia. While some attribute their appearance to north-to-south migration driven by Late Pleistocene climatic cooling events, others emphasize indigenous adaptation to local arid-cool conditions, as seen in the long-standing small flake tool traditions in Southwest China since the late Middle Pleistocene. Notably, the early emergence of Yahuai Cave’s miniaturized lithic artefacts (43 kaBP cal) predates comparable assemblages in northern Jiangxi (26 kaBP), northern Vietnam, and southern Thailand. Geographically, these sites form a descending gradient from mid-altitude mountainous regions (e.g., Guizhou, Guangxi) to low-altitude hilly areas, suggesting a potential southward and eastward diffusion of modern human populations from the hinterland of East Asia. However, the relationship between climatic fluctuations and human behavioural evolution remains complex, requiring further interdisciplinary investigations.

    The discoveries at the Yahuai Cave site underscore that the subtropical-tropical zones of South China and Southeast Asia witnessed the development of complex cultural behaviours among modern humans since 40 kaBP. Detailed technological comparisons of these miniaturized lithic industries will advance our understanding of population dynamics and cultural trajectories during the Late Pleistocene in eastern Eurasia.

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    Lithic technology models and human adaptation behaviors of the Yumidong site in Chongqing
    HE Cunding, LYU Lanxi
    Acta Anthropologica Sinica    2025, 44 (02): 210-219.   DOI: 10.16359/j.1000-3193/AAS.2023.0033
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    Paleolithic archaeological research has predominantly centered on the evolution of lithic technology and the transformation of ancient human subsistence patterns. Scholars worldwide have deliberately correlated the stages of lithic technology evolution with those of human evolution.

    The formation and evolution of lithic technology models are influenced by multiple factors. For an extended period, the prehistoric archaeology community has inclined to employ the perspective of Evolutionary Archaeology to account for the regional variation of culture or lithic industries. However, the role of the Behavioral Ecology perspective in Paleolithic archaeology, particularly in explaining the lithic industries in southern China, has been overlooked. In this paper, under two distinct theoretical frameworks, and by integrating subjective and objective factor analyses, the lithic technology model at the Yumidong Site and the causes of human behavior are systematically elucidated.

    From the perspective of Evolutionary Archaeology, the lithic industry at the Yumidong site, as a regional variant of the main gravel lithic industrial system in southern China, can still be tentatively classified as Lithic Technical Model 1. From a behavioral ecology perspective, the technological landscape of stone products at the Yumidong site is regionally distinctive, differing from the conventional technological patterns hitherto observed. It represents a unique indigenous lithic technological pattern in the Three Gorges region and can be regarded as a new lithic industry or culture.

    We examine and quantify the objective and subjective variables that contributed to the development of the lithic technology type based on a variety of ideas and viewpoints. In terms of raw materials, functional requirements, and passive adaptation, it is posited that the causes of the specificity of the stone tool industry at the Yumidong site are more objective. The role of subjective factors is limited and contingent upon objective factors, and ecological adaptation is the primary determinant of regional cultural characteristics. Nevertheless, subjective technological choices and cultural inheritance are also significant co-factors in the formation and stability of stone tool industrial characteristics. Stone tool technology was highly compatible with the available resources in the environment, and both influenced and converged to facilitate adaptation. This technology and culture system is deeply rooted in the lithic culture of the Three Gorges region. It is a successful instance of ancient humans adapting to the unique environment of the Yangtze River Three Gorges Region and a technological reflection of their distinct survival adaptation behaviors. In southern China, the lithic industry of the Yumidong site is an embodiment of regional technological diversification.

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    Comparison of faunal remains between the Yahuai Cave site and the Bailiandong Cave site, Guangxi
    LIN Minghao, SONG Yanbo, ZHANG Ying, ZHAO Wenya, XIE Guangmao
    Acta Anthropologica Sinica    2025, 44 (03): 499-513.   DOI: 10.16359/j.1000-3193/AAS.2025.0035
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    Guangxi, located in the southwestern part of China, is a crucial region for investigating human origin and evolution from the Palaeolithic to the Neolithic period. During the Upper Palaeolithic, the Last Glacial Maximum and subsequent temperature fluctuations likely posed significant challenges to human subsistence. However, the palaeo-environments and human adaptive strategies in response to local conditions in Guangxi remain largely unexplored.

    This peper focuses on faunal remains excavated from the Yahuai Cave and Bailiandong Cave sites, both spanning a long period from the Upper Palaeolithic to the Neolithic. Based on the modelled radiocarbon dating chronology of the two sites, we investigated human adaptive subsistence behaviours over the past approximately 50,000 years in this region.

    The results show that both the Yahuai Cave and Bailiandong Cave sites were surrounded by low hills, forests, shrubs, and water bodies, which were rich in diverse faunal and floral resources. The appearance of common carps, black carps, and anatids in the Late Upper Palaeolithic period (20,000~12,000 BP) indicates that the water bodies surrounding the sites expanded as the Last Glacial Maximum declined and temperature and precipitation increased.

    Overall, humans at these sites exploited a variety of animals, including mammals, birds, fishes, reptiles, and molluscs. Among them, mammals, especially deer, were the most abundant, highlighting the importance of terrestrial faunal resources to local human groups. Although techniques such as chopping, percussing, cutting, and sawing were used, humans preferred to burn these faunal remains during food preparation. At the Bailiandong Cave site, only two antlers were crafted into tools. In contrast, bone tools made from antlers and shells appeared in all cultural sub-periods at the Yahuai Cave site, indicating the Yahuai Cave community’s continuous need for and utilization of bone tools.

    To estimate the mammalian dietary contribution in different periods at the Yahuai Cave, aurochs and wild boars seemed to contribute the most meat weight, suggesting that local humans employed a “high risk, high return” hunting strategy for subsistence. In the Late Upper Palaeolithic period, humans exploited fishes, molluscs, and other fauna more extensively. Meanwhile, diverse nearby plant resources, including wild rice, were also significantly utilized, indicating the emergence and development of a broad-spectrum subsistence economy. The chronological changes in the type and degree of exploited faunal resources at the two sites provide a valuable case for better understanding human adaptive subsistence in response to fluctuating palaeo-environments in the evolutionary process.

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    Mammalian fauna of the Zhongshan rock shelter site in Bubing Basin, Guangxi
    FAN Yaobin, WANG Wei
    Acta Anthropologica Sinica    2025, 44 (03): 529-544.   DOI: 10.16359/j.1000-3193/AAS.2025.0013
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    The caves in southern China, especially those in the Bubing Basin and Chongzuo region of Guangxi, yield abundant Quaternary mammal fossils. In recent years, a large number of mammal faunas from different Pleistocene periods have been discovered there. These fossil materials have significantly contributed to laying the foundation for understanding the evolution of the Pleistocene mammal fauna. However, due to the scarcity of reports on the late Pleistocene to early Holocene faunas, the evolution of the mammal fauna during this crucial transitional period remains poorly understood.

    Recent research on the Zhongshan rock shelter site in the Bubing Basin, Guangxi, which dates back approximately 14,523~8,472 BP cal, has provided new perspectives on this transitional phase. Based on the detailed identification of mammalian tooth fossils unearthed from the Zhongshan rock - shelter site, the fauna comprises 31 species belonging to 17 families and 6 orders, mainly consisting of medium to large mammals. Among them, Primates and Artiodactyla are the most abundant, reflecting their dominant position in the fauna.

    The fauna from the end of the Pleistocene to the beginning of the Holocene, represented by the Zhongshan rock - shelter site in the Bubing Basin, is mainly characterized by the replacement of extinct species by extant ones. Since the early Pleistocene, the series of cave faunas in the Bubing Basin has shown a gradual decrease in the proportion of extinct species and an increase in extant species. The study of the fauna at the Zhongshan rock shelter site is particularly significant as it fills a critical gap in the biochronological sequence documenting the transition from the Pleistocene fossil fauna to the Holocene modern fauna. This provides a valuable reference for understanding the evolutionary dynamics of Quaternary mammalian faunas in southern China.

    The fossil record highlights the ecological changes in the region during this transitional period, presenting a clearer picture of how mammalian communities adapted to environmental changes. Notably, the richness in the types of primate and deer fossils at the site reflects a diverse ecological landscape dominated by forests. This indicates that the Bubing Basin experienced vegetation recovery after the Last Glacial Maximum, creating a favorable environment for both wildlife and human populations. The presence of biodiversity and abundant resources likely supported human activities in this region.

    In summary, the Zhongshan rock shelter site provides crucial evidence for understanding the late Pleistocene to early Holocene faunal transition in southern China. This research establishes a fundamental framework for further studies on Quaternary mammalian evolution and its environmental context.

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